Spring 2026 Department: Ecology, Evolution, and Environmental Biology

Spring 2026 Ecology, Evolution and Environmental Biology UN1011 BEHAVIOR BIOL-LIVING PRIMATES
Section 001	BEHAVIOR BIOL-LIVING PRIM Call Number: 13083 Points: 3 View Class Schedule & Location in Vergil Enrollment: 22 students (50 max) as of May 18, 2026 Instructor: Marina Cords
Spring 2026 Ecology, Evolution and Environmental Biology UN1111 BEHAVIORAL BIOL-DISC
Section 001	Call Number: 13086 Points: 0 View Class Schedule & Location in Vergil Enrollment: 13 students (25 max) as of May 18, 2026 Instructor: Marina Cords
Section 002	Call Number: 13091 Points: 0 View Class Schedule & Location in Vergil Enrollment: 9 students (25 max) as of May 18, 2026 Instructor: Marina Cords
Spring 2026 Ecology, Evolution and Environmental Biology UN2002 ENVIRONMENTAL BIOLOGY II
Section 001	Call Number: 13096 Points: 4 View Class Schedule & Location in Vergil Enrollment: 17 students (40 max) as of May 18, 2026 Notes: Labs meets W or Th 4:10-7:00. Students choose a lab section Instructor: Matthew I Palmer
Spring 2026 Ecology, Evolution and Environmental Biology UN3087 CONSERVATION BIOLOGY
Section 001	Call Number: 13098 Points: 3 View Class Schedule & Location in Vergil Enrollment: 27 students (30 max) as of May 18, 2026 Instructor: Dustin Partridge
Spring 2026 Ecology, Evolution and Environmental Biology UN3187 CONSERVATION BIOLOGY-DISC
Section 001	Call Number: 13104 Points: 0 View Class Schedule & Location in Vergil Enrollment: 20 students (20 max) as of May 18, 2026 / Full Instructor: Dustin Partridge
Section 002	Call Number: 13105 Points: 0 View Class Schedule & Location in Vergil Enrollment: 7 students (20 max) as of May 18, 2026 Instructor: Dustin Partridge
Spring 2026 Ecology, Evolution and Environmental Biology UN3215 FORENSIC OSTEOLOGY
Section 001	Call Number: 13163 Points: 3 View Class Schedule & Location in Vergil Enrollment: 12 students (15 max) as of May 18, 2026 Notes: EBHS students have priority at first class session Instructor: Jill S Shapiro
Spring 2026 Ecology, Evolution and Environmental Biology UN3320 Biological Interactions in a Changing Wo
Section 001	Bio Interactions-Changing Call Number: 16840 Points: 3 View Class Schedule & Location in Vergil Enrollment: 5 students (20 max) as of May 18, 2026 Notes: Pre-requisites: Environmental Biology I & II or approval by Instructor: Rachel O Cohen
Spring 2026 Ecology, Evolution and Environmental Biology UN3910 THE NEANDERTALS
Section 001	Call Number: 13168 Points: 4 View Class Schedule & Location in Vergil Enrollment: 9 students (12 max) as of May 18, 2026 Instructor: Jill S Shapiro
Spring 2026 Ecology, Evolution and Environmental Biology UN3992 THESIS RESEARCH SEMINAR
Section 001	Call Number: 13188 Points: 3 View Class Schedule & Location in Vergil Enrollment: 9 students (20 max) as of May 18, 2026 Instructors: Matthew I Palmer and Lukas J Musher and Maria Strangas
Spring 2026 Ecology, Evolution and Environmental Biology UN3994 EBHS SENIOR THESIS SEMINAR
Section 001	EBHS SENIOR THESIS SEMINA Call Number: 13195 Points: 3 View Class Schedule & Location in Vergil Enrollment: 2 students (6 max) as of May 18, 2026 Instructor: Jill S Shapiro
Spring 2026 Ecology, Evolution and Environmental Biology UN3998 INDEPENDENT STUDY
Section 001	Call Number: 13206 Points: 1-3 View Class Schedule & Location in Vergil Enrollment: 0 students (6 max) as of May 18, 2026 Instructor: Jill S Shapiro
Section 002	Call Number: 18062 Points: 1-3 View Class Schedule & Location in Vergil Enrollment: 2 students (2 max) as of May 18, 2026 / Full Instructor: Dustin Partridge
Section 003	Call Number: 20201 Points: 1-3 View Class Schedule & Location in Vergil Enrollment: 2 students (2 max) as of May 18, 2026 / Full Instructor: Andres Bendesky
Section 004	Call Number: 20390 Points: 1-3 View Class Schedule & Location in Vergil Enrollment: 0 students (2 max) as of May 18, 2026 Instructor: Marina Cords
Spring 2026 Ecology, Evolution and Environmental Biology GU4055 PRINCIPLES & APPS MOD DNA SEQUENCING TEC
Section 001	PRINCIPLES & APPS MOD DNA Call Number: 13238 Points: 3 View Class Schedule & Location in Vergil Enrollment: 8 students (16 max) as of May 18, 2026 Notes: Introductory biology and/or genetics. Computer programming e Instructor: Andres Bendesky
Spring 2026 Ecology, Evolution and Environmental Biology GU4105 Intermediate Statistics for Ecology and
Section 001	Intermed Stats-Ecology & Call Number: 13242 Points: 3 View Class Schedule & Location in Vergil Enrollment: 12 students (20 max) as of May 18, 2026 Notes: Some prior intro course to R and Stats required, or equivale Instructor: Steffen Foerster
Spring 2026 Ecology, Evolution and Environmental Biology GU4112 ICHTHYOLOGY
Section 001	Call Number: 13243 Points: 3 View Class Schedule & Location in Vergil Enrollment: 16 students (18 max) as of May 18, 2026 Notes: Some classes will be held at AMNH Instructor: Pedro Braganca
Spring 2026 Ecology, Evolution and Environmental Biology GU4126 INTRO TO CONSERVATION GENETICS
Section 001	INTRO TO CONSERVATION GEN Call Number: 13245 Points: 3 View Class Schedule & Location in Vergil Enrollment: 7 students (15 max) as of May 18, 2026 Instructor: Rachel S Welt
Spring 2026 Ecology, Evolution and Environmental Biology GU4135 URBAN ECOLOGY & DESIGN
Section 001	Call Number: 13247 Points: 3 View Class Schedule & Location in Vergil Enrollment: 32 students (45 max) as of May 18, 2026 Notes: Some weeks will have Friday field trips Instructor: Matthew I Palmer
Spring 2026 Ecology, Evolution and Environmental Biology GU4192 INTRO TO LANDSCAPE ANALYSIS
Section 001	INTRO TO LANDSCAPE ANALYS Call Number: 13250 Points: 3 View Class Schedule & Location in Vergil Enrollment: 17 students (25 max) as of May 18, 2026 Instructors: Marcia N Macedo and Jay M Schoen
Spring 2026 Ecology, Evolution and Environmental Biology GU4201 ECO, BEHAVIOR & CONSERVATION OF MAMMALS
Section 001	ECO, BEHAVIOR & CONSERVAT Call Number: 13254 Points: 3 View Class Schedule & Location in Vergil Enrollment: 12 students (12 max) as of May 18, 2026 / Full Instructor: Scott Lapoint
Spring 2026 Ecology, Evolution and Environmental Biology GU4340 HUMAN ADAPTATION
Section 001	Call Number: 16846 Points: 3 View Class Schedule & Location in Vergil Enrollment: 12 students (18 max) as of May 18, 2026 Instructor: Volney K Friedrich
Spring 2026 Ecology, Evolution and Environmental Biology GU4670 Introduction to Geographical Information
Section 001	Introduction to GIS Call Number: 13258 Points: 3 View Class Schedule & Location in Vergil Enrollment: 33 students (34 max) as of May 18, 2026 Instructor: Viorel Popescu
Spring 2026 Ecology, Evolution and Environmental Biology GR5851 THESIS DEVELOPMENT SEMINAR
Section 001	THESIS DEVELOPMENT SEMINA Call Number: 13259 Points: 3 View Class Schedule & Location in Vergil Enrollment: 11 students (20 max) as of May 18, 2026 Instructor: Bekka Brodie
Spring 2026 Ecology, Evolution and Environmental Biology GR6110 FUNDAMENTALS OF EVOLUTION
Section 001	Call Number: 13260 Points: 3 View Class Schedule & Location in Vergil Enrollment: 24 students (30 max) as of May 18, 2026 Instructors: Kira Delmore and Justin Bernstein
Spring 2026 Ecology, Evolution and Environmental Biology GR6300 RESEARCH SEMINAR
Section 001	Call Number: 13261 Points: 1 View Class Schedule & Location in Vergil Enrollment: 37 students (100 max) as of May 18, 2026 Instructor: Maria Diuk-Wasser
Spring 2026 Ecology, Evolution and Environmental Biology GR6905 GRAD SEM IN CONSERVATION BIOL
Section 001	GRAD SEM IN CONSERVATION Call Number: 13262 Points: 3 View Class Schedule & Location in Vergil Enrollment: 17 students (25 max) as of May 18, 2026 Instructor: Bekka Brodie
Spring 2026 Ecology, Evolution and Environmental Biology GR9502 DIRECTED RESEARCH
Section 001	Call Number: 20119 Points: 2-8 View Class Schedule & Location in Vergil Enrollment: 2 students (10 max) as of May 18, 2026 Instructor: Bekka Brodie
Section 002	Call Number: 20210 Points: 2-8 View Class Schedule & Location in Vergil Enrollment: 1 student (3 max) as of May 18, 2026 Instructor: Andres Bendesky
Section 003	Call Number: 20211 Points: 2-8 View Class Schedule & Location in Vergil Enrollment: 1 student (5 max) as of May 18, 2026 Instructor: Duncan Menge
Section 004	Call Number: 20266 Points: 2-8 View Class Schedule & Location in Vergil Enrollment: 3 students (4 max) as of May 18, 2026 Instructor: Viorel Popescu
Section 005	Call Number: 20354 Points: 2-8 View Class Schedule & Location in Vergil Enrollment: 3 students (4 max) as of May 18, 2026 Instructor: Kira Delmore
Section 006	Call Number: 20357 Points: 2-8 View Class Schedule & Location in Vergil Enrollment: 1 student (3 max) as of May 18, 2026 Instructor: Marina Cords
Spring 2026 Ecology, Evolution and Environmental Biology GR9509 DIRECTED READINGS
Section 001	Call Number: 18063 Points: 1-6 View Class Schedule & Location in Vergil Enrollment: 1 student (2 max) as of May 18, 2026 Instructor: Jill S Shapiro
Section 002	Call Number: 20088 Points: 1-6 View Class Schedule & Location in Vergil Enrollment: 3 students (5 max) as of May 18, 2026 Instructor: Viorel Popescu
Section 003	Call Number: 20089 Points: 1-6 View Class Schedule & Location in Vergil Enrollment: 4 students (10 max) as of May 18, 2026 Instructor: Bekka Brodie
Section 004	Call Number: 20264 Points: 1-6 View Class Schedule & Location in Vergil Enrollment: 2 students (3 max) as of May 18, 2026 Instructor: Marcia N Macedo
Section 005	Call Number: 20355 Points: 1-6 View Class Schedule & Location in Vergil Enrollment: 3 students (4 max) as of May 18, 2026 Instructor: Kira Delmore
Section 006	Call Number: 20410 Points: 1-6 View Class Schedule & Location in Vergil Enrollment: 1 student (3 max) as of May 18, 2026 Instructor: Deren Eaton

Spring 2026 Ecology, Evolution and Environmental Biology UN1011
BEHAVIOR BIOL-LIVING PRIMATES

Section 001

Call Number:: 13083
Points:: 3
Enrollment:: 22 students (50 max) as of May 18, 2026
Instructor:: Marina Cords

Spring 2026 Ecology, Evolution and Environmental Biology UN1111
BEHAVIORAL BIOL-DISC

Section 001

Call Number:: 13086
Points:: 0
Enrollment:: 13 students (25 max) as of May 18, 2026
Instructor:: Marina Cords

Section 002

Call Number:: 13091
Points:: 0
Enrollment:: 9 students (25 max) as of May 18, 2026
Instructor:: Marina Cords

Spring 2026 Ecology, Evolution and Environmental Biology UN2002
ENVIRONMENTAL BIOLOGY II

Section 001

Call Number:: 13096
Points:: 4
Enrollment:: 17 students (40 max) as of May 18, 2026
Notes:: Labs meets W or Th 4:10-7:00. Students choose a lab section
Instructor:: Matthew I Palmer

Spring 2026 Ecology, Evolution and Environmental Biology UN3087
CONSERVATION BIOLOGY

Section 001

Call Number:: 13098
Points:: 3
Enrollment:: 27 students (30 max) as of May 18, 2026
Instructor:: Dustin Partridge

Spring 2026 Ecology, Evolution and Environmental Biology UN3187
CONSERVATION BIOLOGY-DISC

Section 001

Call Number:: 13104
Points:: 0
Enrollment:: 20 students (20 max) as of May 18, 2026 / Full
Instructor:: Dustin Partridge

Section 002

Call Number:: 13105
Points:: 0
Enrollment:: 7 students (20 max) as of May 18, 2026
Instructor:: Dustin Partridge

Spring 2026 Ecology, Evolution and Environmental Biology UN3215
FORENSIC OSTEOLOGY

Section 001

Call Number:: 13163
Points:: 3
Enrollment:: 12 students (15 max) as of May 18, 2026
Notes:: EBHS students have priority at first class session
Instructor:: Jill S Shapiro

Spring 2026 Ecology, Evolution and Environmental Biology UN3320
Biological Interactions in a Changing Wo

Section 001

Call Number:: 16840
Points:: 3
Enrollment:: 5 students (20 max) as of May 18, 2026
Notes:: Pre-requisites: Environmental Biology I & II or approval by
Instructor:: Rachel O Cohen

Spring 2026 Ecology, Evolution and Environmental Biology UN3910
THE NEANDERTALS

Section 001

Call Number:: 13168
Points:: 4
Enrollment:: 9 students (12 max) as of May 18, 2026
Instructor:: Jill S Shapiro

Spring 2026 Ecology, Evolution and Environmental Biology UN3992
THESIS RESEARCH SEMINAR

Section 001

Call Number:: 13188
Points:: 3
Enrollment:: 9 students (20 max) as of May 18, 2026
Instructors:: Matthew I Palmer and Lukas J Musher and Maria Strangas

Spring 2026 Ecology, Evolution and Environmental Biology UN3994
EBHS SENIOR THESIS SEMINAR

Section 001

Call Number:: 13195
Points:: 3
Enrollment:: 2 students (6 max) as of May 18, 2026
Instructor:: Jill S Shapiro

Spring 2026 Ecology, Evolution and Environmental Biology UN3998
INDEPENDENT STUDY

Section 001

Call Number:: 13206
Points:: 1-3
Enrollment:: 0 students (6 max) as of May 18, 2026
Instructor:: Jill S Shapiro

Section 002

Call Number:: 18062
Points:: 1-3
Enrollment:: 2 students (2 max) as of May 18, 2026 / Full
Instructor:: Dustin Partridge

Section 003

Call Number:: 20201
Points:: 1-3
Enrollment:: 2 students (2 max) as of May 18, 2026 / Full
Instructor:: Andres Bendesky

Section 004

Call Number:: 20390
Points:: 1-3
Enrollment:: 0 students (2 max) as of May 18, 2026
Instructor:: Marina Cords

Spring 2026 Ecology, Evolution and Environmental Biology GU4055
PRINCIPLES & APPS MOD DNA SEQUENCING TEC

Section 001

Call Number:: 13238
Points:: 3
Enrollment:: 8 students (16 max) as of May 18, 2026
Notes:: Introductory biology and/or genetics. Computer programming e
Instructor:: Andres Bendesky

Spring 2026 Ecology, Evolution and Environmental Biology GU4105
Intermediate Statistics for Ecology and

Section 001

Call Number:: 13242
Points:: 3
Enrollment:: 12 students (20 max) as of May 18, 2026
Notes:: Some prior intro course to R and Stats required, or equivale
Instructor:: Steffen Foerster

Spring 2026 Ecology, Evolution and Environmental Biology GU4112
ICHTHYOLOGY

Section 001

Call Number:: 13243
Points:: 3
Enrollment:: 16 students (18 max) as of May 18, 2026
Notes:: Some classes will be held at AMNH
Instructor:: Pedro Braganca

Spring 2026 Ecology, Evolution and Environmental Biology GU4126
INTRO TO CONSERVATION GENETICS

Section 001

Call Number:: 13245
Points:: 3
Enrollment:: 7 students (15 max) as of May 18, 2026
Instructor:: Rachel S Welt

Spring 2026 Ecology, Evolution and Environmental Biology GU4135
URBAN ECOLOGY & DESIGN

Section 001

Call Number:: 13247
Points:: 3
Enrollment:: 32 students (45 max) as of May 18, 2026
Notes:: Some weeks will have Friday field trips
Instructor:: Matthew I Palmer

Spring 2026 Ecology, Evolution and Environmental Biology GU4192
INTRO TO LANDSCAPE ANALYSIS

Section 001

Call Number:: 13250
Points:: 3
Enrollment:: 17 students (25 max) as of May 18, 2026
Instructors:: Marcia N Macedo and Jay M Schoen

Spring 2026 Ecology, Evolution and Environmental Biology GU4201
ECO, BEHAVIOR & CONSERVATION OF MAMMALS

Section 001

Call Number:: 13254
Points:: 3
Enrollment:: 12 students (12 max) as of May 18, 2026 / Full
Instructor:: Scott Lapoint

Spring 2026 Ecology, Evolution and Environmental Biology GU4340
HUMAN ADAPTATION

Section 001

Call Number:: 16846
Points:: 3
Enrollment:: 12 students (18 max) as of May 18, 2026
Instructor:: Volney K Friedrich

Spring 2026 Ecology, Evolution and Environmental Biology GU4670
Introduction to Geographical Information

Section 001

Call Number:: 13258
Points:: 3
Enrollment:: 33 students (34 max) as of May 18, 2026
Instructor:: Viorel Popescu

Spring 2026 Ecology, Evolution and Environmental Biology GR5851
THESIS DEVELOPMENT SEMINAR

Section 001

Call Number:: 13259
Points:: 3
Enrollment:: 11 students (20 max) as of May 18, 2026
Instructor:: Bekka Brodie

Spring 2026 Ecology, Evolution and Environmental Biology GR6110
FUNDAMENTALS OF EVOLUTION

Section 001

Call Number:: 13260
Points:: 3
Enrollment:: 24 students (30 max) as of May 18, 2026
Instructors:: Kira Delmore and Justin Bernstein

Spring 2026 Ecology, Evolution and Environmental Biology GR6300
RESEARCH SEMINAR

Section 001

Call Number:: 13261
Points:: 1
Enrollment:: 37 students (100 max) as of May 18, 2026
Instructor:: Maria Diuk-Wasser

Spring 2026 Ecology, Evolution and Environmental Biology GR6905
GRAD SEM IN CONSERVATION BIOL

Section 001

Call Number:: 13262
Points:: 3
Enrollment:: 17 students (25 max) as of May 18, 2026
Instructor:: Bekka Brodie

Spring 2026 Ecology, Evolution and Environmental Biology GR9502
DIRECTED RESEARCH

Section 001

Call Number:: 20119
Points:: 2-8
Enrollment:: 2 students (10 max) as of May 18, 2026
Instructor:: Bekka Brodie

Section 002

Call Number:: 20210
Points:: 2-8
Enrollment:: 1 student (3 max) as of May 18, 2026
Instructor:: Andres Bendesky

Section 003

Call Number:: 20211
Points:: 2-8
Enrollment:: 1 student (5 max) as of May 18, 2026
Instructor:: Duncan Menge

Section 004

Call Number:: 20266
Points:: 2-8
Enrollment:: 3 students (4 max) as of May 18, 2026
Instructor:: Viorel Popescu

Section 005

Call Number:: 20354
Points:: 2-8
Enrollment:: 3 students (4 max) as of May 18, 2026
Instructor:: Kira Delmore

Section 006

Call Number:: 20357
Points:: 2-8
Enrollment:: 1 student (3 max) as of May 18, 2026
Instructor:: Marina Cords

Spring 2026 Ecology, Evolution and Environmental Biology GR9509
DIRECTED READINGS

Section 001

Call Number:: 18063
Points:: 1-6
Enrollment:: 1 student (2 max) as of May 18, 2026
Instructor:: Jill S Shapiro

Section 002

Call Number:: 20088
Points:: 1-6
Enrollment:: 3 students (5 max) as of May 18, 2026
Instructor:: Viorel Popescu

Section 003

Call Number:: 20089
Points:: 1-6
Enrollment:: 4 students (10 max) as of May 18, 2026
Instructor:: Bekka Brodie

Section 004

Call Number:: 20264
Points:: 1-6
Enrollment:: 2 students (3 max) as of May 18, 2026
Instructor:: Marcia N Macedo

Section 005

Call Number:: 20355
Points:: 1-6
Enrollment:: 3 students (4 max) as of May 18, 2026
Instructor:: Kira Delmore

Section 006

Call Number:: 20410
Points:: 1-6
Enrollment:: 1 student (3 max) as of May 18, 2026
Instructor:: Deren Eaton

NOTE: Students, Instructors, and Staff should use Vergil Course Search at vergil.columbia.edu/vergil/ for more detailed, accurate, and timely course information.
Class meeting days, times and classroom assignments are now only appearing in Vergil.

Spring 2026 Department: Ecology, Evolution, and Environmental Biology

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BEHAVIOR BIOL-LIVING PRIM

Bio Interactions-Changing

EBHS SENIOR THESIS SEMINA

PRINCIPLES & APPS MOD DNA

Intermed Stats-Ecology &

INTRO TO CONSERVATION GEN

INTRO TO LANDSCAPE ANALYS

ECO, BEHAVIOR & CONSERVAT

Introduction to GIS

THESIS DEVELOPMENT SEMINA

GRAD SEM IN CONSERVATION

NOTE: Students, Instructors, and Staff should use Vergil Course Search at vergil.columbia.edu/vergil/ for more detailed, accurate, and timely course information.Class meeting days, times and classroom assignments are now only appearing in Vergil.

Spring 2026 Department: Ecology, Evolution, and Environmental Biology

For easier printing select theplain text version of this department listingand be sure to print using LANDSCAPE mode

BEHAVIOR BIOL-LIVING PRIM

Bio Interactions-Changing

EBHS SENIOR THESIS SEMINA

PRINCIPLES & APPS MOD DNA

Intermed Stats-Ecology &

INTRO TO CONSERVATION GEN

INTRO TO LANDSCAPE ANALYS

ECO, BEHAVIOR & CONSERVAT

Introduction to GIS

THESIS DEVELOPMENT SEMINA

GRAD SEM IN CONSERVATION

NOTE: Students, Instructors, and Staff should use Vergil Course Search at vergil.columbia.edu/vergil/ for more detailed, accurate, and timely course information.
Class meeting days, times and classroom assignments are now only appearing in Vergil.

For easier printing select the
plain text version of this department listing
and be sure to print using LANDSCAPE mode